Descripcion

Plants scrambling and twining, climbing at least 3 m high, with distinct dichasially branching long and short shoots, shoots bifariously pilose with erect to re- curved trichomes to 400 µm, basally corky, with yel- lowish bark, internodes of long shoots 25–100 mm, ca. 1–2 mm diam.; internodes of short shoots only 5–20 mm, 0.5–1 mm diam. Leaves with channeled petioles 1.5–5 mm, with 0 to 2(to 4) colleters at the adaxial base; blades 5 to 30 (on short shoots rarely more than 15 mm), 4–12 mm wide, lanceolate to ovate, basally rounded, apically acute to acuminate; petioles and blades puberulous to hirsute throughout with recurved trichomes to 100–200 µm; lateral veins indistinct. In- florescences extra-axillary, always 1 per node, sciadioi- dal, 4- to 7-flowered, 2 to 5 flowers open synchronously, sessile; floral bracts ca. 0.5 × 0.3 mm, ovate, abaxially with trichomes; pedicels 0.3–1 mm, uniformly densely hirsute with erect trichomes 200–400 µm. Flowers fra- grant (fide Tweedie s.n. P00214015); flower buds ca. 2.5 × 1.4 mm, conical; calyx campanulate, abaxially with spreading trichomes to 200 µm, with 1 large col- leter in sinuses each; lobes basally fused, ca. 1 × 0.5 mm, ovate-lanceolate, apically acute; corolla cream- colored to yellow; rotate-campanulate (-cupulate), ca. 2.25 × 4.5 mm; lobes ca. 2–2.5 × 0.9 mm, fused to around 1/6 of length, lanceolate, apically obtuse, gla- brous, margins revolute, apically often slightly twisted. Staminal corona cyathiform-cupulate, ca. 0.7–0.8 × 0.9–1.3 mm; lobes fused for about 1/3 of length, rect- angular, somewhat pleated, erect, apically crenulate; forming ovoid pouch in interstaminal position to 0.35 mm deep. Gynostegium cylindrical, ca. 1 × 0.7 mm, sessile; anther wings extending beyond anther proper, at angle of ca. 15°, in same plane as anther; anthers ca. 400 µm, about as long as wide, trapezoidal, abaxially slightly convex, anther wings ca. 275 µm, anther wings of adjacent anthers parallel to each other; connective appendages ca. 400–600 × 200 µm, lanceolate, as cending, margins and apex incurved. Pollinarium with corpusculum ca. 70 × 40 µm, obovoid or ellipsoid; caudicles 40–50 µm, cylindrical, S-shaped, concave c onvex; pollinia subapically attached to caudicles, ca. 100 × 50 µm, ovoid, ovate in cross-section, slightly spreading. Style-head ca. 250 × 350 µm; upper part ca. 175 µm, conical, umbonate. Follicles normally 2 per flower, angle between mericarps ca. 20°, 30–40 mm, narrowly oblong. Seeds ca. 5 mm, oblong (fide Meyer, 1944).