Descripcion

Monoecious shrubs up to 3 m tall; stems erect, un armed, sparsely strigose with sessile glandular tri- chomes; internodes 1.5–7.5 cm. Leaves verticillate, 3 to 4 per node, upright, sometimes patent, subsessile, petioles 0.1–0.3 cm; blade ovate to elliptic, 0.4–6.5 × 0.3–4 cm, apex acute or obtuse, base cuneate, margin entirely crenate, slightly revolute, scabrous adaxially and sparsely to densely strigose abaxially, both sur- faces with sessile glandular trichomes, chartaceous to coriaceous, discolorous, venation pinnate, prominent on abaxial surface. Inflorescence frondose; flowers ar- ranged in lateral spikes, 2 to 3 per leaf axil, rarely sol- itary, capituliform, to 0.5 cm diam., cylindrical in fruit, up to 1 cm, peduncles 2–3 cm, strigose; floral bracts whorled, elliptic, apex acute, strigose with sessile glan- dular trichomes, green, 3.5–4 mm, basal and apical bracts free; calyx tubular, ca. 1.5 mm, margin bifid, strigose, accrescent in fruit; corolla lilac, yellow, or white, 4–5 mm, sparsely strigose. Fruit rounded, 1–2 mm diam., splitting into 2 brown cluses, enclosed by accrescent calyx.

Distribucion y Habitat

Lippia balansae is found in northwestern Argentina (Salta), southern Brazil (Mato Grosso do Sul and Paraná), and central and southern Paraguay. It has been considered endemic to Paraguay (Tron- coso, 1961), but in the present contribution material from Argentina and Brazil was found, thus extending the distribution of the taxon. Although Lippia balansae has a restricted distribu tion in Paraguay, it is very abundant in this country, and many collections have been studied during this work. This species is found in the Humid Chaco and Alto Paraná Atlantic Forest in eastern Paraguay. It gen erally grows on hills in sandy or calcareous soils and has also been collected in flooded areas.

Ejemplares de referencia

Colector N° Colect. Especie Departamento Provincia Imagen
  • Balansa, B.
    		• 1033 Guairá
  • Balansa, B.
    		• 1033 Guairá
  • Balansa, B.
    		• 1033 a (si 66721) Caaguazú
  • Balansa, B.
    		• 1033 a (si 66721) Caaguazú
  • Balansa, B.
    		• 1033 a (si 66722) Caaguazú
  • Basualdo, I.
    		• 1733 Tavaí Caazapá
  • Bordas, E.
    		• 3837 Guairá
  • Burkart, A.
    		• 18640 Cordillera
  • Caballero Mármori, G.
    		• 226 Alto Paraná
  • Degen, R.
    		• 599 Cordillera
  • Degen, R.
    		• 613 Cordillera
  • Fernández Casas, F. J.
    		• 5733 Alto Paraná
  • Fernández Casas, F. J.
    		• 5896 Curuguaty Canindeyú
  • Hahn, W. J.
    		• 882 Caaguazú
  • Hahn, W. J.
    		• 996 Canindeyú
  • Hahn, W. J.
    		• 1377 Canindeyú
  • Harley, R. M.
    		• 28042 Caaguazú
  • Hassler, E.
    		• 4608 Canindeyú
  • Hassler, E.
    		• 6075 Paraguarí
  • Hassler, E.
    		• 6735 Cordillera
  • Hassler, E.
    		• 8653 Guairá
  • Hassler, E.
    		• 12484 Central
  • Jörgensen, P.
    		• 3778 (si 66557) Guairá
  • Jörgensen, P.
    		• 3778 (si 66558) Guairá
  • Jörgensen, P.
    		• 3778 (si 66558) Guairá
  • Krapovickas, A.
    		• 12503 Caaguazú
  • Krapovickas, A.
    		• 13260 Cordillera
  • López, María G.
    		• 254 Alto Paraná
  • Mereles, F.
    		• 7518 Guairá
  • Morrone, O.
    		• 297 Caaguazú
  • Ortiz, M.
    		• 871 Caazapá
  • Pedersen, T. M.
    		• 7572 Cordillera
  • Pedersen, T. M.
    		• 12302 Guairá
  • Pedersen, T. M.
    		• 14663 Mato Grosso do Sul
  • Rojas, T.
    		• 4826 Cordillera
  • Rojas, T.
    		• 4826 a Cordillera
  • Rojas, T.
    		• 4893 Guairá
  • Rojas, T.
    		• 5004 Caaguazú
  • Rojas, T.
    		• 5880 Caazapá
  • Rojas, T.
    		• 9241 Villarrica Guairá
  • Schinini, A.
    		• 1696 Cordillera
  • Schinini, A.
    		• 1965 Cordillera
  • Schinini, A.
    		• 1987 Cordillera
  • Schinini, A.
    		• 1990 Cordillera
  • Schinini, A.
    		• 1991 Eusebio Ayala Cordillera
  • Schinini, A.
    		• 3394 Cordillera
  • Schinini, A.
    		• 9090 San Pedro
  • Schinini, A.
    		• 9132 Caaguazú
  • Schinini, A.
    		• 22932 Caaguazú
  • Schinini, A.
    		• 27392 Alto Paraná
  • Schinini, A.
    		• 27841 Caazapá
  • Schinini, A.
    		• 28102 Caaguazú
  • Schinini, A.
    		• 30205 b Caaguazú
  • Schinini, A.
    		• 33371 Canindeyú
  • Schinini, A.
    		• 36160 Caaguazú
  • Simonis, J. E.
    		• 51 Caaguazú
  • Zardini, E. M.
    		• 2426 Caaguazú
  • Zardini, E. M.
    		• 10485 Caaguazú
  • Zardini, E. M.
    		• 10612 Caaguazú
  • Zardini, E. M.
    		• 10740 Caaguazú
  • Zardini, E. M.
    		• 21992 Cordillera
  • Zardini, E. M.
    		• 25585 Caaguazú
  • Zardini, E. M.
    		• 25696 Caaguazú
  • Zardini, E. M.
    		• 25972 Caaguazú
  • Zardini, E. M.
    		• 26286 Cordillera
  • Zardini, E. M.
    		• 26662 Cordillera
  • Zardini, E. M.
    		• 26938 Tobatí Cordillera
  • Zardini, E. M.
    		• 27471 Cordillera
  • Zardini, E. M.
    		• 38225 Caaguazú

    Nombre Vulgar y Usos

    Tipo y Observaciones

    Material Tipo: Paraguay. Guairá: “Itanga, près de Villarrica,” 17 Feb. 1876, B. Balansa 1033 (lectotipo, G 307903!, designado por F. Mirra et al., Ann. Missouri Bot. Gard. 109: 446. 2024; isolectotipos, P 713663!, P 713664!, P 713665!, SI 3488!).
    Observaciones: Phenology. Lippia balansae flowers and fruits from late September to May.

    Tipo de Lippia calliclada Briq.